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- Apocynaceae
- Cynanchum
- Cynanchum louiseae
Cynanchum louiseae — black swallowwort
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Facts
Black swallowwort, native to Europe, was first noted in 1854 in Ipswich, Massachusetts. Although it still has a fairly restricted distribution outside New England, it can be quite invasive, forming dense stands and crowding out other vegetation.
Habitat
Anthropogenic (man-made or disturbed habitats), forest edges, forests, meadows and fields, shores of rivers or lakes
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Flower petal color
-
- blue to purple
- other
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- opposite: there are two leaves per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Stamen number
- 5
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit length
- 50–70 mm
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower diameter
- 6–8 mm
- Flower number
- 5–7
- Flower petal color
-
- blue to purple
- other
- Flower reproductive parts
- the flower has both pollen- and seed-producing parts
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the flower has two or more completely separate styles
- Fused stamen clusters
- there is one cluster of fused stamens
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Hypanthium length
- 0 mm
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Length of peduncle
- 5–10 mm
- Marks on petals
- there are no noticeable marks on the petals
- Nectar spur
- the flower has no nectar spurs
- Number of branches in umbel
- 0
- Number of carpels
- 2
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Number of styles
- 2
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Perianth shape
- the perianth is rotate (platter-shaped, the corolla flattened, circular, with nearly horizontally spreading lobes)
- Petal and sepal arrangement
- the flower includes three distinct cycles of petal- or sepal-like structures
- Petal and sepal colors
-
- blue to purple
- other
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal folding in bud
- the petals are arranged some other way in bud
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal hairs (Viola)
- NA
- Petal hairs on inner/upper surface
- there are hairs on the inner/upper petal surface
- Petal length
- 1.5–3 mm
- Petal number
- 5
- Petal shape
- the petal outline is roughly triangular
- Petal tips (Cuscuta)
- NA
- Raceme attachment (Veronica)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Scales inside corolla
- no
- Sepal and petal color
- the sepals are different from the petals
- Sepal appearance
- the sepals are green or brown, and leaf-like in texture
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal color
-
- blue to purple
- green to brown
- yellow
- Sepal features
- one or more sepals have glands on the surface of the sepal
- Sepal number
- 5
- Sepal orientation
- the sepals are slightly curved outwards from the corolla
- Sepal shape
- the sepal outline is ovate (widest below the middle and broadly tapering at both ends)
- Sepals fused only to sepals
- the sepals are fused to each other (not other flower parts), at least near their bases
- Stamen appendages
- stamen appendages are present
- Stamen attachment
- the stamens are attached at or near the bases of the petals or tepals
- Stamen number
- 5
- Stamens fused
- the stamens are attached to one another at or near their bases
- Staminodes
- there are no staminodes on the flower
- Style petal-like
- the styles are not petal-like
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule ribs
- NA
- Capsule splitting
- NA
- Carpel beak length
- 0 mm
- Fruit (pyxis) dehiscence
- NA
- Fruit beak length
- 0 mm
- Fruit cross-section
- the fruit is round in cross-section
- Fruit features (Brassicaceae)
- NA
- Fruit length
- 50–70 mm
- Fruit locules
- one
- Fruit shape
- the fruit is ovoid (egg-shaped)
- Fruit type (general)
- the fruit is dry and splits open when ripe
- Fruit type (specific)
- the fruit is a follicle (has one ovary that splits along one side to release the seeds)
- Fruit width
- 8 mm
- Hair type on fruit
- NA
- Hairs on fruit
- the fruits are not hairy
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit does not have a stipe
- Placenta arrangement
- the plant has parietal placentation, where ovules develop on the wall or slight outgrowths of the wall forming broken partitions within a compound ovary
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Seeds comose
- hairs are present
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is milky and opaque, and may be white or colored
- Sap color
- the sap is white
-
Growth form
- Growth form
- the plant is an herb (it has self-supporting stems)
- Lifespan
- the plant lives more than two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
- Underground organs
- the plant has a rhizome (a horizontal underground stem with roots growing from it)
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Hairs on underside of leaf
- the underside of the leaf is fuzzy or hairy
- Hairs on upper side of leaf
- the upper side of the leaf is fuzzy or hairy
- Leaf arrangement
- opposite: there are two leaves per node along the stem
- Leaf blade base
- the leaf has a distinct leaf stalk (petiole)
- Leaf blade base shape
-
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- the base of the leaf blade is rounded
- Leaf blade bloom
- the underside of the leaf has no noticeable bloom
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade length
- 50–70 mm
- Leaf blade primary vein pattern
- the secondary veins branch off at intervals from the primary vein
- Leaf blade shape
- the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf blade vein pattern
- the major veins of the leaf blade branch, but do not rejoin
- Leaf blade width
- 20–30 mm
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf spines
- there are no spines on the leaf edges
- Leaf stalk
- the leaves have leaf stalks
- Leaf stalk attachment to leaf
- the petiole attaches at the basal margin of the leaf blade
- Leaf stalk base
- the petiole base is narrow where it attaches to the stem
- Leaf stalk length
- 10–15 mm
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf tip
- the tip of the leaf blade is acuminate (tapers to a long, thin point)
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf types
- There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
- Leaf variation
- the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- there are two leaves per node along the stem
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule edges
- NA
- Stipule features
- NA
- Stipule fused to leaf stalk
- NA
- Stipule length
- 0 mm
- Stipule shape
- NA
- Stipules
- there are no stipules on the plant
- Teeth per side of leaf blade
- 0
-
Place
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Specific habitat
-
- edges of forests
- forests
- man-made or disturbed habitats
- meadows or fields
- shores of rivers or lakes
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Hair between stem nodes
- the stem has hairs between the nodes
- Hairs between stem nodes
- the hairs on the stem are plain, without glands or branches, and not tangled
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- 40–80 cm
- Stem hair distribution
- the hairs on the stem are distributed more of less uniformly
- Stem orientation
- the stems are upright or angled outwards
- Stem roughness between nodes
- the stem does not feel rough
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
Wetland status
Not classified
In New England
Distribution
- Connecticut
- present, invasive, prohibited
- Maine
- present
- Massachusetts
- present, invasive, prohibited
- New Hampshire
- present, invasive, prohibited
- Rhode Island
- present, invasive
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Massachusetts
- unranked (S-rank: SNR)
From Flora Novae Angliae dichotomous key
1. Cynanchum louiseae Kartesz & Gandhi E
black swallowwort. Cynanchum nigrum (L.) Pers.; Vincetoxicum nigrum (L.) Pers. • CT, MA, ME, NH, RI, VT. Fields, roadsides, forest edges, abandoned homesteads, banks.
Native to North America?
No
Sometimes confused with
- Cynanchum rossicum:
- petals pink to maroon or yellow-white, without hairs, with lobes 2.5-4.5 mm long (vs. C. louisiae, with petals purple-black, minutely hairy on the inner surface, with lobes 1.5-3 mm long).
Synonyms
- Cynanchum nigrum (L.) Pers.
- Vincetoxicum nigrum (L.) Pers.