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• All other flowering non-woody plants
• All other herbaceous, flowering dicots
• Epigaea repens

Epigaea repens — trailing-arbutus

Facts

Trailing-arbutus flowers in early spring. It was designated the State plant of Massachusetts in 1918. The seeds are dispersed by ants that carry the soft placental tissue back to their nests with the seeds attached.

Habitat

Anthropogenic (man-made or disturbed habitats), forest edges, forests

Characteristics

• Clonal plantlets

  Bulbils

  the plant does not appear to have bulbils

  Bulblets replace flowers

  there are no bulblets where the flowers are located

• Flowers

  Anther attachment

  the anther is attached by its base to the filament

  Anther color

  the anthers show no hint of a pink, reddish or purplish tint

  Anther opening

  the anthers have narrow slits or furrows that run lengthwise along the anthers

  Anther spurs

  the anthers do not have spurs on them

  Calyx growth after flowering

  the calyx does not grow to cover or partially cover the fruit

  Calyx symmetry

  there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)

  Carpels fused

  the carpels are fused to one another

  Cilia on petals

  the petal margins do not have cilia

  Cleistogamous flowers

  there are no cleistogamous flowers on the plan

  Corolla morphology

  the tube of the flower is curved upwards

  Corolla palate

  no

  Corona lobe length

  0 mm

  Epicalyx

  the flower does not have an epicalyx

  Epicalyx number of parts

  0

  Filament surface

  • the filament has rough hairs or scales on it
  • the filament is smooth, with no hairs or scales

  Flower appearance

  the flowers appear at the same time as the leaves

  Flower description

  the flower has a superior ovary, and lacks a hypanthium

  Flower number

  2–10

  Flower orientation

  the flower points upwards or is angled outwards

  Flower petal color

  • pink to red
  • white

  Flower reproductive parts

  • the flower has both pollen- and seed-producing parts
  • the flower has either only pollen- or only seed-producing parts

  Flower symmetry

  there are two or more ways to evenly divide the flower (the flower is radially symmetrical)

  Flowers sunken into stem

  no

  Form of style

  the style is unbranched, but it has two or more narrow appendages that are part of the stigma

  Fringed petal edges

  the petals are not fringed

  Fused stamen clusters

  NA

  Fusion of sepals and petals

  • both the petals and sepals are separate and not fused
  • the petals or the sepals are fused into a cup or tube

  Hairs on flower stalk

  the flower stalk has hairs on it

  Horns in hoods (Asclepias)

  NA

  Hypanthium

  the flower does not have a hypanthium

  Hypanthium length

  0 mm

  Inflorescence length

  20–50 mm

  Inflorescence one-sided

  the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks

  Inner tepals (Rumex)

  NA

  Interior flower disk

  the flower does not have an interior disc

  Length of flower stalk

  1–4 mm

  Length of peduncle

  0 mm

  Marks on petals

  • the petals have spots or streaks on them
  • there are no noticeable marks on the petals

  Nectar spur

  the flower has no nectar spurs

  Number of branches in umbel

  0

  Number of carpels

  4–5

  Number of pistils

  1

  Number of sepals, petals or tepals

  there are five petals, sepals, or tepals in the flower

  Number of styles

  1

  Ovary position

  the ovary is above the point of petal and/or sepal attachment

  Perianth shape

  the perianth is salverform (with a slender basal tube and a more-or-less horizontally spreading mouth)

  Petal and sepal arrangement

  the flower includes two cycles of petal- or sepal-like structures

  Petal and sepal colors

  • pink to red
  • white

  Petal appearance

  the petals are thin and delicate, and pigmented (colored other than green or brown)

  Petal base

  the petal narrows gradually or does not narrow at the base

  Petal folds or pleats

  the petals of the flower do not have folds or plaits

  Petal glandular dots or scales

  no

  Petal hairs (Viola)

  NA

  Petal hairs on inner/upper surface

  there are hairs on the inner/upper petal surface

  Petal length

  6–10 mm

  Petal length relative to sepals

  the petals are longer than the sepals

  Petal nectaries

  the petals do not have nectaries

  Petal number

  5

  Petal shape

  • the petal outline is oblong (rectangular, but with rounded ends)
  • the petal outline is ovate (widest below the middle and broadly tapering at both ends)

  Petal tip shape

  the petal tip is obtuse (bluntly pointed)

  Petal tips (Cuscuta)

  NA

  Raceme attachment (Veronica)

  NA

  Reproductive system

  all the flowers on each plant have only carpels or only stamens, with only one type being present on each plant (dioecious)

  Scales inside corolla

  no

  Sepal and petal color

  the sepals are different from the petals

  Sepal appearance

  the sepals are green or brown, and leaf-like in texture

  Sepal appendages

  the sepals do not have appendages on them

  Sepal appendages (Oenothera)

  NA

  Sepal auricles

  the sepals have no auricles

  Sepal color

  • green to brown
  • white

  Sepal features

  NA

  Sepal length

  5–6 mm

  Sepal number

  5

  Sepal relative length

  NA

  Sepal shape

  the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)

  Sepal texture

  the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture

  Sepal tip shape

  the sepal tip is acuminate (tapers to a very narrow point)

  Sepal uniformity

  all the sepals are about the same size

  Sepals fused only to sepals

  the sepals are separate from one another

  Spur length

  0 mm

  Spur number

  NA

  Stamen appendages

  stamen appendages are absent

  Stamen attachment

  the stamens are attached at or near the bases of the petals or tepals

  Stamen lengths differ

  the stamens are all approximately the same length

  Stamen morphology

  the stamens within each cycle are the same

  Stamen number

  10

  Stamen position relative to petals

  NA

  Stamen relative length

  anything

  Stamens fused

  the stamens are not attached to one another

  Staminodes

  • the flower has staminodes
  • there are no staminodes on the flower

  Stigma position

  the stigmas are positioned at the tip of the style

  Style petal-like

  the styles are not petal-like

  Style relative length

  the stigma does not protrude beyond the mouth of the corolla

  Surface of ovary

  the ovary surface is textured with tiny points, bumps or wrinkles

  Umbel flower reproductive parts

  NA

  Upper lip of bilabiate corolla

  NA

• Fruits or seeds

  Achene relative orientation

  NA

  Achene shape

  NA

  Achene surface (Polygonum)

  NA

  Achene type

  NA

  Berry color

  NA

  Capsule color (Viola)

  NA

  Capsule ribs

  the capsule has five prominent ribs or wings

  Capsule splitting

  the capsule splits by five main valves, teeth or pores

  Carpel beak length

  0 mm

  Fruit (pyxis) dehiscence

  NA

  Fruit cross-section

  the fruit is round in cross-section

  Fruit features (Brassicaceae)

  NA

  Fruit locules

  • five
  • three
  • two

  Fruit shape

  • the fruit is flat or strongly compressed
  • the fruit is spherical

  Fruit stalk orientation

  the fruits point upward or spread or curve outward

  Fruit type (general)

  the fruit is fleshy

  Fruit type (specific)

  the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)

  Fruit width

  5–8 mm

  Hair type on fruit

  • the hairs on the fruits appear tangled or woolly
  • the hairs on the fruits have glands at their tips

  Hairs on fruit

  the fruits have hairs on them

  Legumes (Fabaceae)

  NA

  Mericarp length

  0 mm

  Mericarp segment shape (Desmodium)

  NA

  Other markings on berry

  NA

  Ovary stipe

  the ovary or fruit does not have a stipe

  Ovary stipe length

  0 mm

  Placenta arrangement

  the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves

  Rows of seeds in fruit (Brassicaceae)

  NA

  Schizocarpic fruit compression

  NA

  Schizocarpic fruit segments

  0

  Seed length

  Up to 0.5 mm

  Seed number

  At least 100

  Seed relative length

  the seed is about as long as it is wide

  Seed surface

  the seed is covered with reticulate markings (a netlike pattern due to splitting and rejoining of lines or ridges)

  Seeds comose

  no hairs

  Septum in fruit (Brassicaceae)

  NA

  Wings on fruit

  the fruit does not have wings on it

  prickles on fruits

  the fruits do not have thorn-like defensive structures

• Glands or sap

  Glands on leaf blade

  the leaf blades do not have glandular dots or scales

  Sap

  the sap is clear and watery

  Sap color

  the sap is clear

• Growth form

  Growth form

  the plant is a subshrub (small shrub with partially herbaceous stems)

  Horizontal rooting stem

  the plant does not have stolons

  Lifespan

  the plant lives more than two years

  Parasitism

  the plant is not parasitic

  Plant color

  the leaves or young stems of the plant are green

  Plants darken when dry

  no

  Spines on plant

  the plant has no spines

  Underground organs

  the plant has a rhizome (a horizontal underground stem with roots growing from it)

• Leaves

  Bracteole edges

  the bracteole margins are entire (without teeth, lobes or hairs)

  Bracteole number (Apiaceae)

  0

  Bracteoles

  the plant has bracteoles between the primary bracts and the flowers

  Bracts in plantain (Plantago)

  NA

  Final leaf segment length (compound lvs only)

  0 mm

  Final leaf segment length to width ratio (compound lvs only)

  0

  Final leaf segment width (compound lvs only)

  0 mm

  Floral bracts

  the flower has one or more bracts associated with it

  Flower bract length

  4–7 mm

  Hairs on leaf stalk

  the petiole has hairs on it

  Hairs on underside of leaf

  • the underside of the leaf is fuzzy or hairy
  • the underside of the leaf is not hairy, or it has very few hairs

  Hairs on upper side of leaf

  • the upper side of the leaf is fuzzy or hairy
  • the upper side of the leaf is not hairy, or it has very few hairs

  Hooked hairs on underside of leaf

  no

  Inflated hairs on leaf

  the leaf blade does not have inflated hairs on it

  Leaf arrangement

  alternate: there is one leaf per node along the stem

  Leaf blade base

  the leaf has a distinct leaf stalk (petiole)

  Leaf blade base shape

  • the base of the leaf blade is cordate (heart-shaped, has rounded lobes at the base)
  • the base of the leaf blade is rounded

  Leaf blade base symmetry

  the leaf blade base is symmetrical

  Leaf blade bloom

  the underside of the leaf has no noticeable bloom

  Leaf blade edges

  the edge of the leaf blade is entire (has no teeth or lobes)

  Leaf blade flatness

  the leaf is flat (planar) at the edges

  Leaf blade hairs

  • the leaf blade has simple hairs with no glands, and not tangled or wooly
  • the leaf blade has tangled or woolly-looking hairs

  Leaf blade length

  20–100 mm

  Leaf blade primary vein pattern

  the secondary veins branch off at intervals from the primary vein

  Leaf blade shape

  • the leaf blade is elliptic (widest near the middle and tapering at both ends)
  • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)

  Leaf blade surface colors

  the upper side of the leaf blade is relatively uniform in color

  Leaf blade texture

  the leaf blade is coriaceous (has a firm, leathery texture)

  Leaf blade vein pattern

  the major veins form a net-like pattern due to splitting and rejoining

  Leaf blade veins

  the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)

  Leaf blade width

  15–55 mm

  Leaf duration

  the leaves remain green all winter

  Leaf form

  the leaves are green, with an expanded blade and a leaf-like texture

  Leaf hair orientation

  the hairs are standing up straight or curved in different directions

  Leaf shiny

  the upper side of the leaf is dull or slightly shiny

  Leaf spines

  there are no spines on the leaf edges

  Leaf stalk

  the leaves have leaf stalks

  Leaf stalk attachment to leaf

  the petiole attaches at the basal margin of the leaf blade

  Leaf stalk base

  the petiole base is narrow where it attaches to the stem

  Leaf stalk length

  10–50 mm

  Leaf teeth and lobes

  the edge of the leaf blade is entire (has no teeth or lobes)

  Leaf tip

  • the tip of the leaf blade is acute (sharply pointed)
  • the tip of the leaf blade is obtuse (bluntly pointed)

  Leaf tufts in axils

  there are no clusters of smaller leaves growing out of axils

  Leaf type

  the leaves are simple (lobed or unlobed but not separated into leaflets)

  Leaf types

  There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).

  Leaf variation

  the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem

  Leaflet number

  0

  Leaflet petiolules

  NA

  Leaves per node

  there is one leaf per node along the stem

  Pinnately compound leaf type

  NA

  Specific leaf type

  the leaves are simple (lobed or unlobed but not separated into leaflets)

  Stipels

  NA

  Stipule edges

  NA

  Stipule features

  NA

  Stipule fused to leaf stalk

  NA

  Stipule length

  0 mm

  Stipule shape

  NA

  Stipules

  there are no stipules on the plant

  Teeth per side of leaf blade

  0

• Place

  Habitat

  terrestrial

  New England state

  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont

  Specific habitat

  • edges of forests
  • forests
  • man-made or disturbed habitats

• Scent

  Plant odor

  the plant has a pleasant smell, for example anise, fruit, mint or resin

• Stem, shoot, branch

  Branched tendrils

  NA

  Direction of stem hairs

  the hairs point mostly upwards to outwards

  Flowering stem cross-section

  the flowering stem is circular, or with lots of small angles so that it is roughly circular

  Hair between stem nodes

  the stem has hairs between the nodes

  Hooked hairs on stem between nodes

  no

  Leaves on stem

  there is at least one full leaf above the base of the flowering stem

  Plant height

  30–50 cm

  Stem bloom

  there is no powdery or waxy film on the stem

  Stem hair distribution

  the hairs on the stem are distributed more of less uniformly

  Stem nodes swollen

  the stem is not swollen at the nodes

  Stem orientation

  the stems trail at the base, but may turn upwards at the tips

  Stem roughness between nodes

  the stem does not feel rough

  Stem succulence

  the stems are not succulent

  Tendril origin

  NA

  Tendrils

  the plant does not have tendrils

  Wings on stem

  the stem does not have wings on it

Wetland status

Not classified

New England distribution and conservation status

From the dichotomous key of Flora Novae Angliae