Galium tricornutum — rough-fruited bedstraw
![Plant form: Galium tricornutum. ~ By Dr. Nasip Demirkus. ~ Copyright © 2024 Dr. Nasip Demirkus. ~ nasip[at]hotmail.com](https://newfs.s3.amazonaws.com/taxon-images-1000s1000/Rubiaceae/galium-tricornutum-ha-ndemirkus1.jpg)
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Facts
Rough-fruited bedstraw is introduced in New England, where it has been collected only in Rhode Island and northeastern Massachusetts. Its fruits are called "rough" for the many pointed projections that pepper their surfaces (possibly also contributing to the species name, "tricornutum" for "three-horned." The flower stalks (pedicels) are strongly bent backward (recurved) when in flower and fruit.
Habitat
Anthropogenic (man-made or disturbed habitats), meadows and fields
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
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enlargeCharacteristics
- Habitat
- terrestrial
- New England state
- Rhode Island
- Flower petal color
-
- green to brown
- white
- yellow
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- whorled: there are three or more leaves per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
- there are four petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Stamen number
- 4
- Fruit type (general)
- the fruit is dry but does not split open when ripe
- Fruit length
- 0.3–0.5 mm
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Calyx symmetry
- NA
- Carpels fused
- the carpels are fused to one another
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Filament surface
- the filament is smooth, with no hairs or scales
- Flower description
- the flower has an inferior ovary, with or without a hypanthium
- Flower petal color
-
- green to brown
- white
- yellow
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is lobed at the tip, and unbranched
- Fused stamen clusters
- NA
- Fusion of sepals and petals
- the petals or the sepals are fused into a cup or tube
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Nectar spur
- the flower has no nectar spurs
- Number of pistils
- 2
- Number of sepals, petals or tepals
- there are four petals, sepals, or tepals in the flower
- Petal and sepal arrangement
- the flower includes only one cycle of petals or sepals
- Petal and sepal colors
-
- green to brown
- white
- yellow
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal hairs (Viola)
- NA
- Petal number
- 4
- Petal tips (Cuscuta)
- NA
- Reproductive system
- all the flowers have both carpels and stamens (synoecious)
- Scales inside corolla
- no
- Sepal and petal color
- NA
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal number
- 0
- Stamen attachment
- the stamens are not attached to the petals or tepals
- Stamen number
- 4
- Stamen position relative to petals
- the stamens are lined up with the petals
- Staminodes
- there are no staminodes on the flower
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule ribs
- NA
- Capsule splitting
- NA
- Fruit (pyxis) dehiscence
- NA
- Fruit beak length
- 0 mm
- Fruit features (Brassicaceae)
- NA
- Fruit length
- 0.3–0.5 mm
- Fruit locules
- two
- Fruit type (general)
- the fruit is dry but does not split open when ripe
- Fruit type (specific)
- the fruit is a schizocarp (when dry it splits into sections, each holding one or more seeds)
- Legumes (Fabaceae)
- NA
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit does not have a stipe
- Placenta arrangement
- the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- the fruit is not flattened
- Schizocarpic fruit segments
- 2
- Septum in fruit (Brassicaceae)
- NA
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Lifespan
- the plant lives only a single year or less
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Plants darken when dry
- no
- Spines on plant
- the plant has no spines
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Hairs on underside of leaf
-
- the underside of the leaf is fuzzy or hairy
- the underside of the leaf is not hairy, or it has very few hairs
- Leaf arrangement
- whorled: there are three or more leaves per node along the stem
- Leaf blade base
-
- the leaf has a distinct leaf stalk (petiole)
- the leaf has no stalk
- Leaf blade base shape
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade length
- 10–32 mm
- Leaf blade shape
- the leaf blade is oblanceolate (lance-shaped, but with the widest point above the middle of the leaf blade)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf spines
- there are no spines on the leaf edges
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf variation
- the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
-
- there are five leaves per node along the stem
- there are seven or more leaves per node along the stem
- there are six leaves per node along the stem
- Specific leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule features
- NA
- Stipules
-
- the plant has stipules
- there are no stipules on the plant
-
Place
- Habitat
- terrestrial
- New England state
- Rhode Island
- Specific habitat
-
- man-made or disturbed habitats
- meadows or fields
-
Scent
- Plant odor
- the plant does not have much of a smell
-
Stem, shoot, branch
- Branched tendrils
- NA
- Flowering stem cross-section
- the flowering stem is roughly square
- Hair between stem nodes
-
- the stem has hairs between the nodes
- the stem has no hairs between the nodes
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- 5–80 cm
- Stem roughness between nodes
- the stem feels rough due to hairs or prickles
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
Wetland status
Not classified
In New England
Distribution
- Connecticut
- absent
- Maine
- absent
- Massachusetts
- absent
- New Hampshire
- absent
- Rhode Island
- present
- Vermont
- absent
Conservation status
None
From Flora Novae Angliae dichotomous key
19. Galium tricornutum Dandy E
rough-fruited bedstraw. Galium tricorne Stokes, pro parte • RI. Fields, waste areas.
Native to North America?
No
Sometimes confused with
Synonyms
- Galium tricorne Stokes, pro parte