• You are here:
• Dichotomous Key
• Plantaginaceae
• Gratiola
• Gratiola lutea

Gratiola lutea — golden hedge-hyssop

Facts

Despite its name, golden hedge-hyssop is not found in hedges, but is a plant of lake and river shores, mostly on sandy soils. The tubular flowers are usually bright yellow with flairing, irregular lobes.

Habitat

Floodplain (river or stream floodplains), fresh tidal marshes or flats, lacustrine (in lakes or ponds), shores of rivers or lakes

Characteristics

• Clonal plantlets

  Bulbils

  the plant does not appear to have bulbils

  Bulblets replace flowers

  there are no bulblets where the flowers are located

• Flowers

  Anther opening

  the anthers have narrow slits or furrows that run lengthwise along the anthers

  Anther spurs

  the anthers do not have spurs on them

  Calyx symmetry

  there is only one way to evenly divide the calyx (the calyx is bilaterally symmetrical)

  Carpels fused

  the carpels are fused to one another

  Cleistogamous flowers

  there are no cleistogamous flowers on the plan

  Corolla morphology

  the flower has two prominent lips

  Corolla palate

  no

  Corona lobe length

  0 mm

  Epicalyx

  the flower does not have an epicalyx

  Epicalyx number of parts

  0

  Flower description

  the flower has a superior ovary, and lacks a hypanthium

  Flower length

  12–16 mm

  Flower petal color

  yellow

  Flower reproductive parts

  the flower has both pollen- and seed-producing parts

  Flower symmetry

  there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)

  Flowers sunken into stem

  no

  Form of style

  the style is lobed at the tip, and unbranched

  Fused stamen clusters

  NA

  Fusion of sepals and petals

  the petals or the sepals are fused into a cup or tube

  Horns in hoods (Asclepias)

  NA

  Hypanthium

  the flower does not have a hypanthium

  Hypanthium length

  0 mm

  Inflorescence one-sided

  the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks

  Inner tepals (Rumex)

  NA

  Length of flower stalk

  5–15 mm

  Marks on petals

  there are no noticeable marks on the petals

  Nectar spur

  the flower has no nectar spurs

  Number of branches in umbel

  0

  Number of carpels

  2

  Number of pistils

  1

  Number of sepals, petals or tepals

  there are five petals, sepals, or tepals in the flower

  Number of styles

  1

  Ovary position

  the ovary is above the point of petal and/or sepal attachment

  Perianth shape

  • the perianth is campanulate (bell-shaped, with a tube about as long as wide, flaring at the mouth)
  • the perianth is cylindric (tube-shaped)

  Petal and sepal arrangement

  the flower includes two cycles of petal- or sepal-like structures

  Petal and sepal colors

  yellow

  Petal appearance

  the petals are thin and delicate, and pigmented (colored other than green or brown)

  Petal folds or pleats

  the petals of the flower do not have folds or plaits

  Petal hairs (Viola)

  NA

  Petal hairs on inner/upper surface

  there are hairs on the inner/upper petal surface

  Petal length

  12–16 mm

  Petal number

  5

  Petal tips (Cuscuta)

  NA

  Raceme attachment (Veronica)

  NA

  Reproductive system

  all the flowers have both carpels and stamens (synoecious)

  Scales inside corolla

  no

  Sepal and petal color

  the sepals are different from the petals

  Sepal appendages

  the sepals do not have appendages on them

  Sepal appendages (Oenothera)

  NA

  Sepal color

  green to brown

  Sepal length

  4–5 mm

  Sepal number

  5

  Sepal shape

  the sepal outline is lanceolate (lance-shaped; narrow, gradually tapering from the base to the tip)

  Sepal uniformity

  one or more of the sepals is much narrower or shorter than the others

  Sepals fused only to sepals

  the sepals are separate from one another

  Spur length

  0 mm

  Spur number

  NA

  Stamen attachment

  the stamens are attached at or near the bases of the petals or tepals

  Stamen lengths differ

  the stamens are didynamous (two long stamens and two short ones)

  Stamen morphology

  the stamens within a cycle differ in length or width

  Stamen number

  1 or 2

  Stamen relative length

  anything

  Staminodes

  the flower has staminodes

  Style petal-like

  the styles are not petal-like

  Umbel flower reproductive parts

  NA

  Upper lip of bilabiate corolla

  the upper lip of the bilabiate corolla has two lobes

• Fruits or seeds

  Achene relative orientation

  NA

  Achene shape

  NA

  Achene surface (Polygonum)

  NA

  Achene type

  NA

  Berry color

  NA

  Capsule color (Viola)

  NA

  Capsule splitting

  the capsule splits by four main valves, teeth or pores

  Carpel beak length

  0 mm

  Fruit (pyxis) dehiscence

  NA

  Fruit features (Brassicaceae)

  NA

  Fruit length

  2–3 mm

  Fruit length relative to sepals

  the fruit is shorter than its associated sepals

  Fruit locules

  two

  Fruit shape

  the fruit is spherical

  Fruit type (general)

  the fruit is dry and splits open when ripe

  Fruit type (specific)

  the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)

  Hair type on fruit

  NA

  Hairs on fruit

  the fruits are not hairy

  Legumes (Fabaceae)

  NA

  Mericarp length

  0 mm

  Mericarp segment shape (Desmodium)

  NA

  Other markings on berry

  NA

  Ovary stipe

  the ovary or fruit does not have a stipe

  Placenta arrangement

  the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves

  Rows of seeds in fruit (Brassicaceae)

  NA

  Schizocarpic fruit compression

  NA

  Schizocarpic fruit segments

  0

  Septum in fruit (Brassicaceae)

  NA

  Wings on fruit

  the fruit does not have wings on it

  prickles on fruits

  the fruits do not have thorn-like defensive structures

• Glands or sap

  Glands on leaf blade

  the leaf blades have glandular dots or scales

  Sap

  the sap is clear and watery

  Sap color

  the sap is clear

• Growth form

  Growth form

  the plant is an herb (it has self-supporting stems)

  Horizontal rooting stem

  the plant does not have stolons

  Lifespan

  the plant lives more than two years

  Parasitism

  the plant is not parasitic

  Plant color

  the leaves or young stems of the plant are green

  Plants darken when dry

  no

  Spines on plant

  the plant has no spines

  Underground organs

  the plant has a rhizome (a horizontal underground stem with roots growing from it)

• Leaves

  Bracteole number (Apiaceae)

  0

  Bracteoles

  the plant has bracteoles between the primary bracts and the flowers

  Bracts in plantain (Plantago)

  NA

  Final leaf segment length (compound lvs only)

  0 mm

  Final leaf segment length to width ratio (compound lvs only)

  0

  Final leaf segment width (compound lvs only)

  0 mm

  Hairs on leaf stalk

  NA

  Inflated hairs on leaf

  the leaf blade does not have inflated hairs on it

  Leaf arrangement

  opposite: there are two leaves per node along the stem

  Leaf blade base

  the leaf has no stalk

  Leaf blade base symmetry

  the leaf blade base is symmetrical

  Leaf blade bloom

  the underside of the leaf has no noticeable bloom

  Leaf blade edges

  • the edge of the leaf blade has teeth
  • the edge of the leaf blade is entire (has no teeth or lobes)

  Leaf blade flatness

  the leaf is flat (planar) at the edges

  Leaf blade length

  8–30 mm

  Leaf blade primary vein pattern

  the secondary veins branch off at intervals from the primary vein

  Leaf blade shape

  • the leaf blade is lanceolate (lance-shaped; widest below the middle and tapering at both ends)
  • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)

  Leaf blade surface colors

  the upper side of the leaf blade is relatively uniform in color

  Leaf blade texture

  the leaf blade has a fleshy or spongy texture

  Leaf blade vein pattern

  the major veins of the leaf blade branch, but do not rejoin

  Leaf blade veins

  the leaf blade has one main vein running from the base towards the tip (it may or may not have secondary veins)

  Leaf blade width

  3–7 mm

  Leaf duration

  the leaves drop off in winter (or they whither but persist on the plant)

  Leaf form

  the leaves are green, with an expanded blade and a leaf-like texture

  Leaf sheath length

  0 mm

  Leaf shiny

  the upper side of the leaf is dull or slightly shiny

  Leaf spines

  there are no spines on the leaf edges

  Leaf stalk

  the leaves have no leaf stalks, but attach directly to the stem

  Leaf stalk attachment to leaf

  NA

  Leaf stalk base

  NA

  Leaf stalk length

  0 mm

  Leaf teeth and lobes

  • the edge of the leaf blade is entire (has no teeth or lobes)
  • the leaf blade margin has outward-pointing teeth

  Leaf type

  the leaves are simple (lobed or unlobed but not separated into leaflets)

  Leaf types

  There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).

  Leaf variation

  the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem

  Leaflet number

  0

  Leaflet petiolules

  NA

  Leaves per node

  there are two leaves per node along the stem

  Pinnately compound leaf type

  NA

  Specific leaf type

  the leaves are simple (lobed or unlobed but not separated into leaflets)

  Stipels

  NA

  Stipule edges

  NA

  Stipule features

  NA

  Stipule fused to leaf stalk

  NA

  Stipule length

  0 mm

  Stipule shape

  NA

  Stipules

  there are no stipules on the plant

  Teeth per side of leaf blade

  At least 0

• Place

  Habitat

  • aquatic
  • wetlands

  New England state

  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont

  Specific habitat

  • fresh tidal marshes or flats
  • in lakes or ponds
  • river or stream floodplains
  • shores of rivers or lakes

• Scent

  Plant odor

  the plant does not have much of a smell

• Stem, shoot, branch

  Branched tendrils

  NA

  Flowering stem cross-section

  • the flowering stem is circular, or with lots of small angles so that it is roughly circular
  • the flowering stem is roughly square

  Flowering stem width

  1–2.5 mm

  Hair between stem nodes

  • the stem has hairs between the nodes
  • the stem has no hairs between the nodes

  Hairs between stem nodes

  • at least some of the hairs on the stem have glands
  • the stem has no hairs between the nodes

  Hooked hairs on stem between nodes

  no

  Leaves on stem

  there is at least one full leaf above the base of the flowering stem

  Plant height

  10–30 cm

  Stem bloom

  there is no powdery or waxy film on the stem

  Stem orientation

  • the stems are upright or angled outwards
  • the stems trail at the base, but may turn upwards at the tips

  Stem roughness between nodes

  the stem does not feel rough

  Stem succulence

  the stems are succulent

  Tendril origin

  NA

  Tendrils

  the plant does not have tendrils

  Wings on stem

  the stem does not have wings on it

Wetland status

Occurs only in wetlands. (Wetland indicator code: OBL)

New England distribution and conservation status

From the dichotomous key of Flora Novae Angliae