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Plantago arenaria — sand plantain

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Facts

Sand plantain is native to Europe and is a source of the mucilaginous seeds (psyllium) widely used as a laxative. In New England, sand plantain is a weed of disturbed sites.

Habitat

Anthropogenic (man-made or disturbed habitats)

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
terrestrial
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Vermont
Flower petal color
  • green to brown
  • pink to red
Leaf type
the leaves are simple (lobed or unlobed but not separated into leaflets)
Leaf arrangement
opposite: there are two leaves per node along the stem
Leaf blade edges
the edge of the leaf blade is entire (has no teeth or lobes)
Flower symmetry
there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
Number of sepals, petals or tepals
there are four petals, sepals, or tepals in the flower
Fusion of sepals and petals
the petals or the sepals are fused into a cup or tube
Stamen number
4
Fruit type (general)
the fruit is dry and splits open when ripe
Fruit length
3–4 mm
Show all characteristics

  • Clonal plantlets

    Bulbils
    the plant does not appear to have bulbils
    Bulblets replace flowers
    there are no bulblets where the flowers are located

  • Flowers

    Anther color
    the anthers show no hint of a pink, reddish or purplish tint
    Anther opening
    the anthers have narrow slits or furrows that run lengthwise along the anthers
    Anther spurs
    the anthers do not have spurs on them
    Calyx symmetry
    there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
    Carpels fused
    the carpels are fused to one another
    Corolla palate
    no
    Corona lobe length
    0 mm
    Epicalyx
    the flower does not have an epicalyx
    Epicalyx number of parts
    0
    Filament surface
    the filament is smooth, with no hairs or scales
    Flower description
    the flower has a superior ovary, and lacks a hypanthium
    Flower petal color
    • green to brown
    • pink to red
    Flower reproductive parts
    the flower has both pollen- and seed-producing parts
    Flower symmetry
    there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
    Flowers sunken into stem
    no
    Form of style
    the style is lobed at the tip, and unbranched
    Fused stamen clusters
    NA
    Fusion of sepals and petals
    the petals or the sepals are fused into a cup or tube
    Horns in hoods (Asclepias)
    NA
    Hypanthium
    the flower does not have a hypanthium
    Inflorescence length
    5–15 mm
    Inflorescence one-sided
    the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
    Inflorescence width
    10 mm
    Inner tepals (Rumex)
    NA
    Length of peduncle
    20–80 mm
    Marks on petals
    there are no noticeable marks on the petals
    Nectar spur
    the flower has no nectar spurs
    Number of branches in umbel
    0
    Number of sepals, petals or tepals
    there are four petals, sepals, or tepals in the flower
    Number of styles
    1
    Ovary position
    the ovary is above the point of petal and/or sepal attachment
    Perianth shape
    • the perianth is cylindric (tube-shaped)
    • the perianth is salverform (with a slender basal tube and a more-or-less horizontally spreading mouth)
    Petal and sepal arrangement
    the flower includes two cycles of petal- or sepal-like structures
    Petal and sepal colors
    • green to brown
    • pink to red
    Petal appearance
    the petals are green and/or leafy in texture
    Petal folds or pleats
    the petals of the flower do not have folds or plaits
    Petal hairs (Viola)
    NA
    Petal number
    4
    Petal shape
    the petal outline is ovate (widest below the middle and broadly tapering at both ends)
    Petal tips (Cuscuta)
    NA
    Raceme attachment (Veronica)
    NA
    Reproductive system
    all the flowers have both carpels and stamens (synoecious)
    Scales inside corolla
    no
    Sepal and petal color
    the sepals are different from the petals
    Sepal appendages
    the sepals do not have appendages on them
    Sepal appendages (Oenothera)
    NA
    Sepal color
    • green to brown
    • other
    Sepal length
    3 mm
    Sepal number
    4
    Sepal texture
    • the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
    • the sepals are thin and dry, paperlike
    Sepal tip shape
    • the sepal tip is acute (is sharply pointed)
    • the sepal tip is rounded
    Stamen attachment
    the stamens are attached at or near the bases of the petals or tepals
    Stamen number
    4
    Stamen position relative to petals
    the stamens are lined up with the petals
    Stamen relative length
    anything
    Staminodes
    there are no staminodes on the flower
    Style petal-like
    the styles are not petal-like
    Style relative length
    the stigma protrudes beyond the mouth of the corolla
    Umbel flower reproductive parts
    NA
    Upper lip of bilabiate corolla
    NA

  • Fruits or seeds

    Achene relative orientation
    NA
    Achene shape
    NA
    Achene surface (Polygonum)
    NA
    Achene type
    NA
    Berry color
    NA
    Capsule color (Viola)
    NA
    Capsule ribs
    NA
    Capsule splitting
    NA
    Carpel beak length
    0 mm
    Fruit (pyxis) dehiscence
    the pyxis splits on or near the middle
    Fruit features (Brassicaceae)
    NA
    Fruit length
    3–4 mm
    Fruit locules
    two
    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a pyxis (when dry it splits around the middle, and the top falls off, exposing the seeds)
    Legumes (Fabaceae)
    NA
    Mericarp length
    0 mm
    Mericarp segment shape (Desmodium)
    NA
    Other markings on berry
    NA
    Ovary stipe
    the ovary or fruit does not have a stipe
    Placenta arrangement
    the plant has axile placentation, in which the ovules are attached where the septa of a compound ovary are united, usually on the central axis, or to the septa themselves
    Rows of seeds in fruit (Brassicaceae)
    NA
    Schizocarpic fruit compression
    NA
    Schizocarpic fruit segments
    0
    Seed length
    2–3 mm
    Seed number
    1–2
    Seed relative length
    the seed is longer than it is wide
    Septum in fruit (Brassicaceae)
    NA
    Wings on fruit
    the fruit does not have wings on it
    prickles on fruits
    the fruits do not have thorn-like defensive structures

  • Glands or sap

    Glands on leaf blade
    the leaf blades do not have glandular dots or scales
    Sap
    the sap is clear and watery
    Sap color
    the sap is clear

  • Growth form

    Growth form
    the plant is an herb (it has self-supporting stems)
    Horizontal rooting stem
    the plant does not have stolons
    Lifespan
    the plant lives only a single year or less
    Parasitism
    the plant is not parasitic
    Plant color
    the leaves or young stems of the plant are green
    Plants darken when dry
    no
    Spines on plant
    the plant has no spines
    Underground organs
    there is a thickened taproot on the plant

  • Leaves

    Bracteole number (Apiaceae)
    0
    Bracts in plantain (Plantago)
    the bracts are hairy
    Final leaf segment length (compound lvs only)
    0 mm
    Final leaf segment length to width ratio (compound lvs only)
    0
    Final leaf segment width (compound lvs only)
    0 mm
    Floral bracts
    the flower has one or more bracts associated with it
    Hooked hairs on underside of leaf
    no
    Inflated hairs on leaf
    the leaf blade does not have inflated hairs on it
    Leaf arrangement
    opposite: there are two leaves per node along the stem
    Leaf blade base symmetry
    the leaf blade base is symmetrical
    Leaf blade bloom
    the underside of the leaf has no noticeable bloom
    Leaf blade edges
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf blade length
    20–80 mm
    Leaf blade primary vein pattern
    the major veins radiate out from one point at the base
    Leaf blade shape
    the leaf blade is linear (very narrow with more or less parallel sides)
    Leaf blade surface colors
    the upper side of the leaf blade is relatively uniform in color
    Leaf blade texture
    the leaf blade is herbaceous (has a leafy texture)
    Leaf blade width
    1–3 mm
    Leaf duration
    the leaves drop off in winter (or they whither but persist on the plant)
    Leaf form
    the leaves are green, with an expanded blade and a leaf-like texture
    Leaf shiny
    the upper side of the leaf is dull or slightly shiny
    Leaf spines
    there are no spines on the leaf edges
    Leaf stalk
    • the leaves have leaf stalks
    • the leaves have no leaf stalks, but attach directly to the stem
    Leaf stalk attachment to leaf
    • NA
    • the petiole attaches at the basal margin of the leaf blade
    Leaf teeth and lobes
    the edge of the leaf blade is entire (has no teeth or lobes)
    Leaf type
    the leaves are simple (lobed or unlobed but not separated into leaflets)
    Leaf variation
    the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
    Leaflet number
    0
    Leaflet petiolules
    NA
    Leaves per node
    there are two leaves per node along the stem
    Pinnately compound leaf type
    NA
    Specific leaf type
    the leaves are simple (lobed or unlobed but not separated into leaflets)
    Stipels
    NA
    Stipule edges
    NA
    Stipule features
    NA
    Stipule fused to leaf stalk
    NA
    Stipule length
    0 mm
    Stipule shape
    NA
    Stipules
    there are no stipules on the plant
    Teeth per side of leaf blade
    0

  • Place

    Habitat
    terrestrial
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Vermont
    Specific habitat
    man-made or disturbed habitats

  • Scent

    Plant odor
    the plant does not have much of a smell

  • Stem, shoot, branch

    Branched tendrils
    NA
    Direction of stem hairs
    the hairs point mostly upwards to outwards
    Hair between stem nodes
    the stem has hairs between the nodes
    Hairs between stem nodes
    the hairs on the stem are plain, without glands or branches, and not tangled
    Hooked hairs on stem between nodes
    no
    Leaves on stem
    there is at least one full leaf above the base of the flowering stem
    Plant height
    10–60 cm
    Stem bloom
    there is no powdery or waxy film on the stem
    Stem hair distribution
    the hairs on the stem are distributed more of less uniformly
    Stem orientation
    the stems are upright or angled outwards
    Stem roughness between nodes
    the stem does not feel rough
    Tendril origin
    NA
    Tendrils
    the plant does not have tendrils
    Wings on stem
    the stem does not have wings on it

Wetland status

Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)

In New England

Distribution

Connecticut
absent
Maine
absent
Massachusetts
absent
New Hampshire
absent
Rhode Island
absent
Vermont
absent

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
unranked (S-rank: SNR)

From Flora Novae Angliae dichotomous key

1.  Plantago arenaria Waldst. & Kit. E

sand plantain. Plantago indica L., nom. illeg.; P. psyllium L., nom. ambig.; P. ramosa Aschers. 
• CT, MA, ME, NH, VT. Roadsides, parking lots, railroads, waste areas.

Native to North America?

No

Synonyms

  • Plantago indica L., an illegitimate name
  • Plantago psyllium L., an ambiguous name
  • Plantago ramosa Aschers.

Family

Plantaginaceae

Genus

Plantago