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- Geocaulon lividum
Geocaulon lividum — false toadflax
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Facts
False toadflax is a rare plant of northern and high-altitude areas of New England in low-nutrient soils. It also occurs in coastal peatlands of downeast Maine. The single Vermont population, in a tiny bog on Mount Mansfield, has not been seen since 1901. It is the only member of the genus in New England, and one of only two genera in the small family, Comandraceae. It produces small, greenish flowers without petals, which mature into large orange-red fruits that are edible (though not very tasty).
Habitat
Alpine or subalpine zones, bogs, fens (calcium-rich wetlands), mountain summits and plateaus
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
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Characteristics
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Maine
- New Hampshire
- Vermont
- Flower petal color
-
- blue to purple
- green to brown
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
-
- there are five petals, sepals, or tepals in the flower
- there are four petals, sepals, or tepals in the flower
- Fusion of sepals and petals
- both the petals and sepals are separate and not fused
- Stamen number
-
- 4
- 5
- Fruit type (general)
- the fruit is fleshy
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Anther spurs
- the anthers do not have spurs on them
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla palate
- no
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Flower description
- the flower has an inferior ovary, with or without a hypanthium
- Flower petal color
-
- blue to purple
- green to brown
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is knob-like at the tip, and unbranched
- Fusion of sepals and petals
- both the petals and sepals are separate and not fused
- Horns in hoods (Asclepias)
- NA
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Nectar spur
- the flower has no nectar spurs
- Number of pistils
- 1
- Number of sepals, petals or tepals
-
- there are five petals, sepals, or tepals in the flower
- there are four petals, sepals, or tepals in the flower
- Ovary position
- the ovary is below the point of petal and/or sepal attachment
- Petal and sepal arrangement
- the flower includes only one cycle of petals or sepals
- Petal and sepal colors
-
- blue to purple
- green to brown
- Petal appearance
- the petals are thin and delicate, and pigmented (colored other than green or brown)
- Petal folds or pleats
- the petals of the flower do not have folds or plaits
- Petal hairs (Viola)
- NA
- Petal number
- 4–5
- Petal tips (Cuscuta)
- NA
- Reproductive system
-
- all the flowers have both carpels and stamens (synoecious)
- the plant has flowers with both carpels and stamens, as well as flowers with only carpels or only stamens (polygamous)
- Scales inside corolla
- no
- Sepal and petal color
- the petals and sepals are similar
- Sepal appendages
-
- NA
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Stamen attachment
- the stamens are attached at or near the bases of the petals or tepals
- Stamen number
-
- 4
- 5
- Stamen position relative to petals
- the stamens are lined up with the sepals
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene relative orientation
- NA
- Achene shape
- NA
- Achene surface (Polygonum)
- NA
- Achene type
- NA
- Berry color
-
- NA
- orange
- red
- yellow
- Capsule color (Viola)
- NA
- Capsule ribs
- NA
- Capsule splitting
- NA
- Fruit (pyxis) dehiscence
- NA
- Fruit features (Brassicaceae)
- NA
- Fruit locules
- one
- Fruit type (general)
- the fruit is fleshy
- Fruit type (specific)
- the fruit is a drupe (fleshy, with a firm inner ovary wall that encloses a single seed)
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
-
- NA
- the ripe berries are mostly one color without spots or streaks
- Ovary stipe
- the ovary or fruit does not have a stipe
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades do not have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Horizontal rooting stem
- the plant does not have stolons
- Lifespan
- the plant lives more than two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Spines on plant
- the plant has no spines
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Hairs on underside of leaf
- the underside of the leaf is not hairy, or it has very few hairs
- Leaf arrangement
- alternate: there is one leaf per node along the stem
- Leaf blade base
- the leaf has a distinct leaf stalk (petiole)
- Leaf blade base shape
- the base of the leaf blade is cuneate (wedge-shaped, tapers to the base with relatively straight, converging edges), or narrow
- Leaf blade edges
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf blade length
- 10–40 mm
- Leaf blade shape
-
- the leaf blade is elliptic (widest near the middle and tapering at both ends)
- the leaf blade is oblong (rectangular but with rounded ends)
- the leaf blade is obovate (egg-shaped, but with the widest point above the middle of the leaf blade)
- the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf spines
- there are no spines on the leaf edges
- Leaf teeth and lobes
- the edge of the leaf blade is entire (has no teeth or lobes)
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf types
-
- NA
- There is a gradual change in appearance of the leaves from the base (or near the base) of the plant to those from further up on the stem, with leaves progressively changing as one moves higher on the stem (often becoming shorter, or less toothed/lobed, and/or with shorter petioles).
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
- there is one leaf per node along the stem
- Specific leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Stipule features
- NA
- Stipules
- there are no stipules on the plant
-
Place
- Habitat
-
- terrestrial
- wetlands
- New England state
-
- Maine
- New Hampshire
- Vermont
- Specific habitat
-
- alpine or subalpine zones
- bogs
- fens (calcium-rich wetlands)
- mountain summits and plateaus
-
Stem, shoot, branch
- Branched tendrils
- NA
- Flowering stem cross-section
- the flowering stem is circular, or with lots of small angles so that it is roughly circular
- Hairs between stem nodes
- the stem has no hairs between the nodes
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- 7–30 cm
- Stem roughness between nodes
- the stem does not feel rough
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
Wetland status
Occurs in wetlands or non-wetlands. (Wetland indicator code: FAC)
In New England
Distribution
- Connecticut
- absent
- Maine
- present
- Massachusetts
- absent
- New Hampshire
- present
- Rhode Island
- absent
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Maine
- uncommon (S-rank: S3), special concern (code: SC)
- New Hampshire
- extremely rare (S-rank: S1), endangered (code: E)
- Vermont
- extirpated (S-rank: SX)
From Flora Novae Angliae dichotomous key
1. Geocaulon lividum (Richards.) Fern. NC
false toadflax. Comandra lividum Richards. • ME, NH, VT. Subalpine heaths, coastal peatlands, mossy openings in boreal and alpine, evergreen forests. Found mainly in northern and/or high elevation areas of New England, Geocaulon lividum occurs also in coastal peatlands of downeast ME.
Native to North America?
Yes
Sometimes confused with
Synonyms
- Comandra lividum Richards.