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- Humulus
- Humulus lupulus
Humulus lupulus — common hop, hops
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Facts
The female fruit of the common hop is used to flavor beer and prevent bacterial contamination during fermenting. This vine can grow up to thirty feet but dies back each year to its roots.
Habitat
Anthropogenic (man-made or disturbed habitats), floodplain (river or stream floodplains), forests, shrublands or thickets
New England distribution
Adapted from BONAP data
Native: indigenous.
Non-native: introduced (intentionally or unintentionally); has become naturalized.
County documented: documented to exist in the county by evidence (herbarium specimen, photograph). Also covers those considered historical (not seen in 20 years).
State documented: documented to exist in the state, but not documented to a county within the state. Also covers those considered historical (not seen in 20 years).
Note: when native and non-native populations both exist in a county, only native status is shown on the map.
Found this plant? Take a photo and post a sighting.
Characteristics
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Flower petal color
- NA
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf arrangement
-
- alternate: there is one leaf per node along the stem
- opposite: there are two leaves per node along the stem
- Leaf blade edges
-
- the edge of the leaf blade has lobes, or it has both teeth and lobes
- the edge of the leaf blade has teeth
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Fusion of sepals and petals
-
- both the petals and sepals are separate and not fused
- the petals or the sepals are fused into a cup or tube
- Stamen number
- 5
- Fruit type (general)
- the fruit is dry but does not split open when ripe
-
Clonal plantlets
- Bulbils
- the plant does not appear to have bulbils
- Bulblets replace flowers
- there are no bulblets where the flowers are located
-
Flowers
- Anther attachment
- the anther is attached by its base to the filament
- Anther opening
- the anthers have narrow slits or furrows that run lengthwise along the anthers
- Anther spurs
- the anthers do not have spurs on them
- Calyx symmetry
- there are two or more ways to evenly divide the calyx (the calyx is radially symmetrical)
- Carpels fused
- the carpels are fused to one another
- Cleistogamous flowers
- there are no cleistogamous flowers on the plan
- Corolla morphology
- NA
- Corolla palate
- NA
- Corona lobe length
- 0 mm
- Epicalyx
- the flower does not have an epicalyx
- Epicalyx number of parts
- 0
- Flower description
- the flower has a superior ovary, and lacks a hypanthium
- Flower number
- 10–200
- Flower petal color
- NA
- Flower reproductive parts
- the flower has either only pollen- or only seed-producing parts
- Flower symmetry
- there are two or more ways to evenly divide the flower (the flower is radially symmetrical)
- Flowers sunken into stem
- no
- Form of style
- the style is unbranched, but it has two or more narrow appendages that are part of the stigma
- Fringed petal edges
- NA
- Fusion of sepals and petals
-
- both the petals and sepals are separate and not fused
- the petals or the sepals are fused into a cup or tube
- Horns in hoods (Asclepias)
- NA
- Hypanthium
- the flower does not have a hypanthium
- Hypanthium length
- 0 mm
- Inflorescence length
- 10–20 mm
- Inflorescence one-sided
- the flowers are arrayed in a spiral around the inflorescence axis or branches, or occur singly, or in several ranks
- Inner tepals (Rumex)
- NA
- Interior flower disk
- the flower does not have an interior disc
- Length of peduncle
- 7–25 mm
- Marks on petals
- NA
- Nectar spur
- the flower has no nectar spurs
- Number of branches in umbel
- 0
- Number of carpels
- 2
- Number of pistils
- 1
- Number of sepals, petals or tepals
- there are five petals, sepals, or tepals in the flower
- Number of styles
- 1
- Ovary position
- the ovary is above the point of petal and/or sepal attachment
- Petal and sepal arrangement
- the flower includes only one cycle of petals or sepals
- Petal and sepal colors
- green to brown
- Petal appearance
- NA
- Petal base
- NA
- Petal folds or pleats
- NA
- Petal glandular dots or scales
- NA
- Petal hairs (Viola)
- NA
- Petal hairs on inner/upper surface
- NA
- Petal length
- 0 mm
- Petal nectaries
- NA
- Petal number
- 0
- Petal shape
- NA
- Petal tip shape
- NA
- Petal tips (Cuscuta)
- NA
- Petal width
- 0 mm
- Raceme attachment (Veronica)
- NA
- Reproductive system
- all the flowers on each plant have only carpels or only stamens, with only one type being present on each plant (dioecious)
- Scales inside corolla
- NA
- Sepal and petal color
- NA
- Sepal appearance
- the sepals are green or brown, and leaf-like in texture
- Sepal appendages
- the sepals do not have appendages on them
- Sepal appendages (Oenothera)
- NA
- Sepal color
-
- green to brown
- white
- Sepal number
- 5
- Sepal texture
- the sepals are either very thin but flexible, like a membrane, or they are leaf-like in texture
- Sepals fused only to sepals
-
- the sepals are fused to each other (not other flower parts), at least near their bases
- the sepals are separate from one another
- Stamen attachment
-
- NA
- the stamens are not attached to the petals or tepals
- Stamen number
- 5
- Stamen position relative to petals
- the stamens are lined up with the sepals
- Staminodes
- there are no staminodes on the flower
- Style petal-like
- the styles are not petal-like
- Umbel flower reproductive parts
- NA
- Upper lip of bilabiate corolla
- NA
-
Fruits or seeds
- Achene shape
- the achenes are oval in outline
- Achene surface (Polygonum)
- NA
- Achene type
- the fruit is an achene (dry, indehiscent, and usually one-seeded)
- Berry color
- NA
- Capsule color (Viola)
- NA
- Capsule ribs
- NA
- Capsule splitting
- NA
- Carpel beak length
- 0 mm
- Fruit (pyxis) dehiscence
- NA
- Fruit beak length
- 0 mm
- Fruit cross-section
- the fruit is at least somewhat flattened
- Fruit features (Brassicaceae)
- NA
- Fruit length relative to sepals
- the fruit is shorter than its associated sepals
- Fruit locules
- one
- Fruit shape
- the fruit is ovoid (egg-shaped)
- Fruit stalk orientation
- the fruits curve or droop downwards
- Fruit type (general)
- the fruit is dry but does not split open when ripe
- Fruit type (specific)
- the fruit is an achene (dry, usually one-seeded, does not separate or split open at maturity)
- Legumes (Fabaceae)
- NA
- Mericarp length
- 0 mm
- Mericarp segment shape (Desmodium)
- NA
- Other markings on berry
- NA
- Ovary stipe
- the ovary or fruit does not have a stipe
- Placenta arrangement
- the plant has apical placentation, where one or a few ovules develop at the top of a simple or compound ovary
- Rows of seeds in fruit (Brassicaceae)
- NA
- Schizocarpic fruit compression
- NA
- Schizocarpic fruit segments
- 0
- Septum in fruit (Brassicaceae)
- NA
- Wings on fruit
- the fruit does not have wings on it
- prickles on fruits
- the fruits do not have thorn-like defensive structures
-
Glands or sap
- Glands on leaf blade
- the leaf blades have glandular dots or scales
- Sap
- the sap is clear and watery
- Sap color
- the sap is clear
-
Growth form
- Growth form
-
- the plant is a vine (it cannot support its own weight)
- the plant is an herb (it has self-supporting stems)
- Lifespan
- the plant lives more than two years
- Parasitism
- the plant is not parasitic
- Plant color
- the leaves or young stems of the plant are green
- Spines on plant
- there are spines on the plant
- Underground organs
- the plant has a rhizome (a horizontal underground stem with roots growing from it)
-
Leaves
- Bracteole number (Apiaceae)
- 0
- Bracteoles
- the plant has bracteoles between the primary bracts and the flowers
- Bracts in plantain (Plantago)
- NA
- Final leaf segment length (compound lvs only)
- 0 mm
- Final leaf segment length to width ratio (compound lvs only)
- 0
- Final leaf segment width (compound lvs only)
- 0 mm
- Floral bracts
- the flower has one or more bracts associated with it
- Hairs on leaf stalk
- the petiole has hairs on it
- Hairs on underside of leaf
-
- the underside of the leaf is fuzzy or hairy
- the underside of the leaf is not hairy, or it has very few hairs
- Hairs on upper side of leaf
-
- the upper side of the leaf is fuzzy or hairy
- the upper side of the leaf is not hairy, or it has very few hairs
- Hooked hairs on underside of leaf
- no
- Inflated hairs on leaf
- the leaf blade does not have inflated hairs on it
- Leaf arrangement
-
- alternate: there is one leaf per node along the stem
- opposite: there are two leaves per node along the stem
- Leaf blade base
- the leaf has a distinct leaf stalk (petiole)
- Leaf blade base shape
- the base of the leaf blade is cordate (heart-shaped, has rounded lobes at the base)
- Leaf blade base symmetry
- the leaf blade base is asymmetrical
- Leaf blade bloom
- the underside of the leaf has no noticeable bloom
- Leaf blade edges
-
- the edge of the leaf blade has lobes, or it has both teeth and lobes
- the edge of the leaf blade has teeth
- Leaf blade hairs
-
- the leaf blade has hairs with glands at their tips
- the leaf blade has simple hairs with no glands, and not tangled or wooly
- Leaf blade length
- 30–150 mm
- Leaf blade primary vein pattern
- the major veins radiate out from one point at the base
- Leaf blade shape
- the leaf blade is cordate (heart-shaped with backward-facing rounded lobes), or sagittate (arrow-shaped with backward-facing pointed lobes)
- Leaf blade surface colors
- the upper side of the leaf blade is relatively uniform in color
- Leaf blade texture
- the leaf blade is herbaceous (has a leafy texture)
- Leaf blade vein pattern
- the major veins of the leaf blade branch, but do not rejoin
- Leaf duration
- the leaves drop off in winter (or they whither but persist on the plant)
- Leaf form
- the leaves are green, with an expanded blade and a leaf-like texture
- Leaf shiny
- the upper side of the leaf is dull or slightly shiny
- Leaf spines
- there are no spines on the leaf edges
- Leaf stalk
- the leaves have leaf stalks
- Leaf stalk attachment to leaf
- the petiole attaches at the basal margin of the leaf blade
- Leaf stalk length
- Up to 70 mm
- Leaf teeth and lobes
- the leaf has lobes that radiate from the base, somewhat like a hand
- Leaf tip
- the tip of the leaf blade is acuminate (tapers to a long, thin point)
- Leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Leaf types
- NA
- Leaf variation
- the leaves are nearly similar in size, prominence of teeth, and length of stalks throughout the stem
- Leaflet number
- 0
- Leaflet petiolules
- NA
- Leaves per node
-
- there are two leaves per node along the stem
- there is one leaf per node along the stem
- Pinnately compound leaf type
- NA
- Specific leaf type
- the leaves are simple (lobed or unlobed but not separated into leaflets)
- Stipels
- NA
- Stipule features
- NA
- Stipule shape
- the stipules are triangular
- Stipules
- the plant has stipules
-
Place
- Habitat
- terrestrial
- New England state
-
- Connecticut
- Maine
- Massachusetts
- New Hampshire
- Rhode Island
- Vermont
- Specific habitat
-
- forests
- man-made or disturbed habitats
- river or stream floodplains
- shrublands or thickets
-
Scent
- Plant odor
- the plant has a pleasant smell, for example anise, fruit, mint or resin
-
Stem, shoot, branch
- Branched tendrils
- NA
- Direction of stem hairs
- the hairs point downwards, or they bend outwards and then downwards
- Hair between stem nodes
- the stem has hairs between the nodes
- Hairs between stem nodes
-
- at least some of the hairs on the stem are branched
- at least some of the hairs on the stem have glands
- Hooked hairs on stem between nodes
- no
- Leaves on stem
- there is at least one full leaf above the base of the flowering stem
- Plant height
- 100–700 cm
- Stem bloom
- there is no powdery or waxy film on the stem
- Stem hair distribution
- the hairs on the stem are mainly found at the nodes
- Stem orientation
- the stems are upright or angled outwards
- Stem roughness between nodes
- the stem does not feel rough
- Tendril origin
- NA
- Tendrils
- the plant does not have tendrils
- Wings on stem
- the stem does not have wings on it
Wetland status
Usually occurs in non-wetlands, but occasionally in wetlands. (Wetland indicator code: FACU)
In New England
Distribution
- Connecticut
- present
- Maine
- present
- Massachusetts
- present
- New Hampshire
- present
- Rhode Island
- present
- Vermont
- present
Conservation status
Exact status definitions can vary from state to state. For details, please check with your state.
- Massachusetts
- unranked (S-rank: SNR)
var. lupuloides
- Massachusetts
- unranked (S-rank: SNR)
var. lupulus
- Massachusetts
- unranked (S-rank: SNR)
Subspecies and varieties
Humulus lupulus L. ssp. lupulus is known from CT, MA, ME, NH, VT and is non-native, often found in human-disturbed habitats. H. lupulus ssp. americanus (Nutt.) A. & D. Löve is known from CT, MA, ME, NH, RI, VT and is native to New England, where it is found in riparian habitats.
From Flora Novae Angliae dichotomous key
2. Humulus lupulus L. n
common hop. 2a. Humulus americanus Nutt.; H. lupulus L. var. lupuloides E. Small • CT, MA, ME, NH, RI, VT. Riparian forests and shrub thickets, abandoned homesteads, roadsides, waste areas.
1a. Abaxial leaf midrib with 20–100 hairs per cm; abaxial leaf surface with more than 25 glands per 10 mm2 … 2a. H. lupulus ssp. americanus (Nutt.) A. & D. Löve
1b. Abaxial leaf midrib with fewer than 20 hairs per cm; abaxial leaf surface with fewer than 25 glands per 10 mm2 … 2b. H. lupulus ssp. lupulus
Supspecies americanus is known from CT, MA, ME, NH, RI, VT, and is native to New England, usually occurring in riparian habitats. Subspecies lupulus is known from CT, MA, ME, NH, VT and is non-native, often occurring in human-disturbed habitats.
Native to North America?
Yes and no (some introduced)
Sometimes confused with
- Humulus japonicus:
- larger leaf blades with 5-9 lobes separated by narrow sinuses and lacking glandular dots and fruiting spikes with bracts that do not conceal the fruits (vs. H. lupulus, with the larger leaf blades with 3-5 lobes separated by open sinuses and with tiny glandular dots and fruiting spikes with bracts that conceal the fruits).
Synonyms
- Humulus americanus Nutt.
- Humulus lupulus var. lupuloides E. Small