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Utricularia geminiscapa — mixed bladderwort

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Facts

Mixed bladderwort is confined to Northeastern North America, and recently naturalized in New Zealand. Its common name refers to the fact that it produces two types of flowers, those that open and can exchange pollen with other plants (chasmogamous) and those that remain closed, transferring its own pollen to the stigma inside (cleistogamous). Both types of flower can create viable seeds.

Habitat

Bogs, fens (calcium-rich wetlands), lacustrine (in lakes or ponds), shores of rivers or lakes

New England distribution

Adapted from BONAP data

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North America distribution

Adapted from BONAP data

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Characteristics

Habitat
  • aquatic
  • wetlands
New England state
  • Connecticut
  • Maine
  • Massachusetts
  • New Hampshire
  • Rhode Island
  • Vermont
Leaf position
  • some of the leaves are floating at the surface of the water
  • the leaves are all submerged underwater
Leaf arrangement
alternate: there is one leaf per node along the stem
Leaf blade length
10–20 mm
Petal or sepal number
  • there are five petals, sepals, or tepals in the flower
  • there are two petals, sepals, or tepals in the flower
Petal color
yellow
Specific leaf type
the leaf is divided into two equal parts which in turn are divided into two further parts; each of these parts are divided and each of these may be further divided
Floating leaf shape
  • the leaf blade is orbicular (roughly circular, as wide as long)
  • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
Underwater leaf blade width
10–35 mm
Fruit type (general)
the fruit is dry and splits open when ripe
Underwater leaf length
10–20 mm
Show all characteristics
  • Flowers

    Anther color
    there is a noticeable pink, reddish or purplish tint to the anthers
    Carpels fused
    the carpels are fused to one another
    Flower lower lip length
    0–8 mm
    Flower number
    1–8
    Flower position
    • the flowers are above the surface of the water
    • the flowers are below the surface of the water
    Flower symmetry
    there is only one way to evenly divide the flower (the flower is bilaterally symmetrical)
    Inflorescence type
    • the inflorescence has only one flower on it
    • the inflorescence is a raceme (a long unbranched stem with stalked flowers growing along it)
    Length of flower stalk
    5–15 mm
    Length of peduncle
    5–250 mm
    Nectar spur
    the flower has nectar spurs
    Number of carpels
    2
    Ovary position
    the sepals and/or petals are attached below the ovary
    Palate on corolla
    • no
    • yes
    Petal and sepal arrangement
    • the flower includes only one cycle of petals or sepals
    • the flower includes two cycles of petal- or sepal-like structures
    Petal appearance
    the petals are thin and delicate, and pigmented (colored other than green or brown)
    Petal color
    yellow
    Petal fringed edges
    the petals are not fringed
    Petal fusion
    the perianth parts are fused to form a tube, cup, or bell shape
    Petal hairs on inner/upper surface
    there are no hairs on the inner/upper petal surface
    Petal length
    5–8 mm
    Petal number
    0–5
    Petal or sepal number
    • there are five petals, sepals, or tepals in the flower
    • there are two petals, sepals, or tepals in the flower
    Pistil number
    1
    Sepal appearance
    the sepals resemble leaves in color and texture
    Sepal number
    2
    Sepals fused only to sepals
    the sepals are separate from one another
    Stamen number
    1 or 2
    Stamen position relative to petals
    NA
    Stamens fused
    the stamens are not fused to one another
    Stamens fused to petals
    the stamens are fused near the bases of the petals or tepals
    Style length
    At least 0 mm
    Style number
    0–1
  • Fruits or seeds

    Fruit type (general)
    the fruit is dry and splits open when ripe
    Fruit type (specific)
    the fruit is a capsule (splits along two or more seams, apical teeth or pores when dry, to release two or more seeds)
  • Glands or sap

    Oil glands on nodes
    none of the nodes have oil glands
    Sap
    the sap is clear and watery
  • Growth form

    Lifespan
    • the plant lives more than two years
    • the plant lives only a single year or less
    Root septa
    the roots do not have transverse septa
    Roots floating in water
    there are no clusters of roots floating in the water
    Turions
    the plant has turions
    Underground organs
    NA
  • Leaves

    Bract position (Sparganium)
    NA
    Bract relative length
    At least 1379 mm
    Bracts
    the flowers or their pedicels have bracts at their bases
    Floating leaf basal lobes
    no
    Floating leaf blade width
    10–35 mm
    Floating leaf length
    0 mm
    Floating leaf shape
    • the leaf blade is orbicular (roughly circular, as wide as long)
    • the leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    Floating leaf tip
    the tip of the floating leaf blade is rounded, with no point
    Floral bract form
    the bracts are much more lobed, or much less lobed, than the foliage leaves
    Floral bract length
    2–3 mm
    Leaf arrangement
    alternate: there is one leaf per node along the stem
    Leaf blade length
    10–20 mm
    Leaf blade veins
    the lateral veins radiate from the base and continue to spread away from the centerline of the leaf, or branch off the central vein at intervals
    Leaf blade width
    10–35 mm
    Leaf position
    • some of the leaves are floating at the surface of the water
    • the leaves are all submerged underwater
    Leaf special features
    the leaves have bladder-like traps
    Leaf-like branch segments
    0
    Leaf-like branch shape
    the leaf-like branches are round
    Specific leaf type
    the leaf is divided into two equal parts which in turn are divided into two further parts; each of these parts are divided and each of these may be further divided
    Staminate bract edge (Myriophyllum)
    NA
    Stipule appearance
    NA
    Stipule fused to leaf
    NA
    Stipules
    there are no stipules on the plant
    Stipules fused around stem
    NA
    Trap-bladder length
    0.5–5 mm
    Underwater leaf air passage number
    At least 1439
    Underwater leaf air passage relative width
    At least 4336
    Underwater leaf air passage row number
    0
    Underwater leaf blade edges
    the underwater leaf has smooth edges, without teeth
    Underwater leaf blade shape
    • NA
    • the underwater leaf blade is orbicular (roughly circular, as wide as long)
    • the underwater leaf blade is ovate (widest below the middle and broadly tapering at both ends)
    Underwater leaf blade veins
    1–3
    Underwater leaf blade width
    10–35 mm
    Underwater leaf length
    10–20 mm
    Underwater leaf stalk
    yes
    Underwater leaf tip shape
    the tip of the underwater leaf is rounded, with no point
    Veins in floating leaf
    1–3
  • Place

    Habitat
    • aquatic
    • wetlands
    New England state
    • Connecticut
    • Maine
    • Massachusetts
    • New Hampshire
    • Rhode Island
    • Vermont
    Specific habitat
    • bogs
    • fens (calcium-rich wetlands)
    • in lakes or ponds
    • shores of rivers or lakes
  • Stem, shoot, branch

    Flowering stem growth form
    the flowering stem is upright

Wetland status

Occurs only in wetlands. (Wetland indicator code: OBL)

In New England

Distribution

Connecticut
present
Maine
present
Massachusetts
present
New Hampshire
present
Rhode Island
present
Vermont
present

Conservation status

Exact status definitions can vary from state to state. For details, please check with your state.

Massachusetts
unranked (S-rank: SNR)
Rhode Island
extremely rare (S-rank: S1), concern (code: C)
Vermont
uncommon (S-rank: S3)

From Flora Novae Angliae dichotomous key

2.  Utricularia geminiscapa Benj. N

mixed bladderwort. Utricularia clandestina Nutt. ex Gray • CT, MA, ME, NH, RI, VT. Shallow water of lakes, ponds, and peatland pools. This species can be confused with smaller plants of Utricularia macrorhiza. It has ultimate branch segments with 0–2 (–4) latera setae and its smaller turions lack hairs (vs. (0–) 2–4 (–8) lateral setae on ultimate segments and larger turions with abundant hairs in U. macrorhiza).

Native to North America?

Yes

Sometimes confused with

Utricularia vulgaris:
flowers of one type--open-pollinated ones that are elevated above the surface of the water, and winter buds 5.5-13 mm thick (vs. U. geminiscapa, with flowers of two types--open-pollinated ones that are elevated above the surface of the water and closed-pollinated ones that are held beneath the water, and winter buds mostly 2-5 mm thick).

Synonyms

  • Utricularia clandestina Nutt. ex Gray

Genus

Utricularia